The rounded long [ǭ] survived in Primitive Danian, as well as developing form primitive long [ā], but became unrounded [ō] in East Danian and the diphthong [ua] in Ossiriandic (PE19/25). It reappeared later in Ossiriandic as a development from the diphthong [au], but later still reduced to the unrounded long [ō].
References ✧ PE19/25
|Dan. [no change]||
||✧ PE19/25 (ǭ > †ǭ)|
||✧ PE19/25 (ā > †ǭ)|
||✧ PE19/25 (†ǭ > uo > ūa); PE19/25 (†ǭ > uo > ūa)|
||✧ PE19/25 (†ǭ > ō); PE19/25 (†ǭ > ō)|